Resumen
In this study, the structure and spatial patterns of adjacent beech (Fagus sylvatica L.) forest stands, both those having undergone intensive management and those without management intervention over the last 50 years were analyzed. Data was collected from six 1ha permanent research plots. Four plots were established in varying parts of the reserve, comprising entirely of even-aged managed forest stands; two 1-ha permanent plots were located in a non-interventional stand. For each woody stem, basic dendrometrical data and coordinates were measured. The total volume of dead wood (standing and fallen) and its classes of decay were recorded. The structure and species diversity of forest stands were analyzed using skewness and the coefficient of variation of dbh distribution, the Gini index, the Shannon index, the mingling index, the Pielou index of segregation and the Hopkins-Skellam, Pielou-Mountford, Clark-Evans aggregation indices, the diameter differentiation index and Ripley K function. The diameter distribution of the managed stands was generally normal, albeit with a large diameter range and with a typical flat diameter curve. A general lack of giant trees, lower rates of coarse woody debris (from 0.2 to 0.5% of the total living volume) and random tree distribution were also confirmed. Within the investigated area, the diameter differentiation and Gini indices along with the degree of skewness and the coefficient of variation made it possible to monitor the shift from even-aged forest management to spontaneous forest development. In general, spontaneous development was marked mainly by aggregated tree distribution in the middle and lower layer, and a shift from normal to reversed J-shaped dbh distribution. The total volume of deadwood in one of the unmanaged plots amounted to 48.10 m3/ha (9.5% of living wood volume). The use of adequate structural and spatial analysis as indicators of natural forest development are discussed.